Part 1 Genetic variation: genetic and molecular essentials; types of polymorphisms; organization of genetic variation; inbreeding. Part 2 The causes of evolution: mutation; migration; natural selection; random genetic drift. Part 3 Molecular population genetics: molecular polymorphisms; patterns of change in nucleotide and amino acid sequences; polymorphism and divergence; molecular phylogenetics; transposable elements. Part 4 The genetic architecture of complex traits: types of complex traits; phenotypic variation; genetics and environment; artificial selection; correlation between relatives; quantitative genetics of natural populations; complex traits with discrete expression; quantitative trait loci (QTLs).